“Not having oxygen” doesn’t necessarily lead to pain, at least not for many animals. When you hold your breath, the pain you feel isn’t because of a lack of oxygen, but instead from a buildup of carbon dioxide in your lungs (which is actually toxic). The result of this is that asphyxiation by some compounds (particularly water or nitrogen) is actually not particularly painful.
Possibly temperature, in the winter the air can be much colder than the water itself, in the summer imagine being pulled from 20°c water onto hot sand or being held in your hand which could be 30°c, its not pleasant for the fish either way!!
Ps if you are going to eat them, kill them immediately. Do not let them suffer, flipping about being scared out of their minds. Whack them in the head with a blunt object. I do fish for food but if i was a fish i would rather be killed straight away than being left in a bucket or bag, slowly suffocating. Its not nice, put them out.
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